Basically any USN/USMC F/A-18 pilot will go from a Cessna 172 or a Piper through a civi flight school at the beginning of their time at Naval Aviation Schools Command, or on their own, previous to the Navy. After that they'll go into a T-6 Texan II for a few months, then if they get selected for Jets, they'll fly the T-45 Goshawk for they carrier qualification and basic stuff, then if they get Hornets they will get sent to the RAG to learn theyre airframe, in this case, the Hornet.
So they really aren't going into an 'advanced' trainer, its basic stuff in the T-45, then BAM, hornet.
I could be mistaken on a couple points here since I wasn't in the pilot pipeline, rather the NFO pipeline.